Gravely injured, Frostpaw places her trust in a young warrior, convincing him to protect her as she follows StarClan's cryptic directions toward the roots of RiverClan, where she hopes to find the answers she so desperately seeks while staving off a war between the clans.

"As the water buffalo prepare for their yearly migration, a terrible accident leaves their leader unable to go on--and in his place, young Echo is chosen by the Great Spirit to lead his herd. But the older buffalo can't believe such a young animal has been chosen--and refuse to follow his lead, threatening the migration that is necessary for the Great Spirit to bring the rains to the land. Meanwhile, Stride the cheetah is keeping secrets from his...

Firestar's three grandchildren, Hollypaw, Lionpaw, and Jaypaw, struggle to restore faith in StarClan as Sol works to tear apart the bonds between Clans and turn them away from their warrior ancestors.